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Goto:

Clossiana Reuss, 1920

Clossiana Reuss, 1920; Ent. Mitt. 9: 192 nota, TS: Papilio selene Denis & Schiffermüller

= Boloria Moore, [1900]; [Wahlberg]

Clossiana, NSG Voucher Specimen [Wahlberg]
Clossiana (Nymphalidae) ; [BRU2, 43]

700x504 (~51Kb) USA: Corral Pass region (47°00'15N 121°29'02W), Pierce Co., WA, 18.7.2000, Photo © Markku Savela

The exact identification of this species is still unknown, but tentatively assumed to belong into this group.

Clossiana selene ([Schiffermüller], 1775)

. See [About maps] Niittyhopeatäplä Small Pearl-bordered Fritillary Silver-bordered Frtillary Silver Meadow Fritillary Brunfläckig pärlemorfjäril Ängspärlefjäril Braunfleckiger Perlmutterfalter Le Petit Collier Argenté Perlada castaña Dostojka selene
Papilio selene [Schiffermüller], 1775; Ankündung eines systematischen Werkes von den Schmetterlingen der Wienergegend: 321, TL: Austria
euphrosyne (Bergsträsser, 1779)
euphrasia (Lewin, 1795)
silene (Haworth, 1803)
Boloria selene, NSG Voucher Specimen [Wahlberg]
Boloria selene, Butterflies and Moths of North America [BAMONA]
Boloria selene, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Boloria selene, Svenska fjärilar [SVF]
Boloria selene, Norges Sommerfugler [NORLEP]
Clossiana selene, Moths and Butterflies of Europe [MBE]
Clossiana selene, Europaische Schmetterlinge [Mario Maier]
Clossiana selene, Captain's European Butterfly Guide [Simon Coombes]
Clossiana selene ; [NACL, #4464]; [BOW: pl. 2, f. 2-3]; [EBW]; [BARU, #188]; [BRU2: 45, pl. 32, f. 1-6]
Boloria selene ; [BNA: 318, #239, pl. 3, 5, 24, 26]; [BOC, 281]; [Opler]; [Otakar Kudrna]

450x697 (~67Kb) upperside	458x713 (~67Kb) underside	425x374 (~35Kb) upperside
401x353 (~34Kb) underside	500x391 (~75Kb) upperside Haldon woods, Devon, UK 21/05/95, Photo © Simon Coombes	500x389 (~57Kb) underside Haldon woods, Devon, UK 21/05/95, Photo © Simon Coombes
837x1028 (~120Kb) underside Finland, Kuhmalahti, Mattilankylä, Setälänlahti, 23.6.2005, Photo © Harri Arkkio	1371x992 (~199Kb) underside Finland, Kuhmalahti, Mattilankylä, Setälänlahti, 23.6.2005, Photo © Harri Arkkio

Larva on Viola, V. palustris, V. canina, V. riviniana [SPRK], Fragaria, Vaccinium uliginosum [BRU2, 46], Viola nephrophylla, V. glabella [BNA, 318]

C. s. selene

EU, W.Siberia, N.Kazakhstan, Altai, Sayan, Transbaikalia Clossiana selene selene ; [BRU2, 45]

C. s. thalia (Hübner, 1790)

Arctic Europe, N.Siberia ?Papilio thalia Hübner, [1799-1800]; Samml. eur. Schmett. [1] : pl. 11 (1799-1800), f. 57-58
hela (Staudinger, 1861)
obscurum (Sedych, 1977)
Clossiana selene thalia ; [BARU]; [BRU2, 45]

C. s. dilutior (Fixsen, 1887)

E.Amur, Ussuri sugitanii (Seok, 1938)
Clossiana selene dilutior ; [BARU]; [BRU2, 45]

C. s. chibiana (Matsumura, 1927)

Sakhalin, Kunashir Clossiana selene chibiana ; [BARU]; [BRU2, 45]

C. s. myrina (Cramer, [1777])

Papilio myrina Cramer, [1777]; Uitl. Kapellen 2 (9-16): 141, pl. 189, f. B, C
Papilio myrinus Herbst, 1798
Argynnis myrissa Godart, 1819; 253
Brenthis myrina nubes Scudder, 1889; 595, TL: Sanbornton, New Hampshire
ab. nivea Gunder, 1928; Can. Ent. 60: (162-168) (albino), TL: Staten Island, New York
Brenthis selene marilandica Clark, 1941; 384, TL: Beltsville, Maryland
Boloria selene myrina, Butterflies of America [BOA]

Argynnis myrina ; Scudder, 1863, 166; Scudder, 1868, 379; Edwards, 1876, 161 Brenthis myrina ab. nubes ; McDunnough, 1938, 16

Brenthis myrina ; Dyar, 1900, 487; Cary, 1901, 307; Dyar, 1903, Bull. U.S. natn. Mus., 52: 15; Cary, 1906, 439; Elrod, 1906, 69; Franzen, 1914, 366; McDunnough, 1928, 274; Holland, 1931, The Butterfly Book (1931 edn): 105; Clark, 1932, 102; dos Passos & Grey, 1934, 190; McDunnough, 1938, 15; Davenport & Dethier, 1938, 158; Leussler, 1938, 216; Rysgaard, 1939, 193; Comstock, 1940, 53; Brooks, 1942, 34 Brenthis selene myrina ; Clark, 1941, 384 Boloria myrina ; dos Passos & Grey, 1945, 4 Boloria selene marilandica ; Klots, 1951, 89; dos Passos, 1964, 89 Clossiana selene myrina ; [NACL, #4464a]; [BNA, #239] Boloria selene myrina ; Klots, 1951, 89; dos Passos, 1964, 89; Shepard, 1975, in Howe, 246; Kohler, 1977, J. Lep. Soc. 31 (4): 245; [Jonathan Pelham] Boloria myrina ; [Joseph Belicek]

There is split of opinions whether 'myrina' is considered as own species or as ssp. of 'selene'. I have no idea of the "correct" ranking, but the setup here requires me to choose either one or the other, the choice is random, tentative, and with no taxonomic statement assocated with it! -- msa

C. s. nebraskensis (Holland, 1928)

E.Nebraska - Ohio Brenthis myrina var. nebraskensis Holland, 1928; 36, TL: Dodge Co., Nebraska
Boloria selene nebraskensis, Butterflies of America [BOA]

Brenthis myrina var. nebraskensis ; Holland, 1931, The Butterfly Book (1931 edn): 105, pl. LV, f. 8 (type) Brenthis myrina r. nebraskensis ; Leussler, 1938, 216 Brenthis myrina nebraskensis ; McDunnough, 1938, 16 Brenthis selene nebraskensis ; Clark, 1941, 384

Clossiana selene nebraskensis ; [NACL, #4464b] Boloria selene nebrascensis ; Forbes, 1960, 160 (missp.) Boloria selene nebraskensis ; Klots, 1951, 89; Brown, 1957, 64; dos Passos, 1964, 89; Shepard, 1975, in Howe, 246; Kohler, 1977, J. Lep. Soc. 31 (4): 248, f. 2, 6; [BNA, #239]

C. s. sabulocollis (Kohler, 1977)

Nebraska, South Dakota, North Dakota, Colorado Boloria selene sabulocollis Kohler, 1977; J. Lep. Soc. 31 (4): 252, f. 3, 6-7, TL: Smith Lake, Sheridan Co., Nebraska
Boloria selene sabulocollis, Butterflies of America [BOA]
Clossiana selene sabulicollis[sic?] ; [NACL, #4464c]

C. s. tollandensis (Barnes & Benjamin, 1925)

Rocky Mountains (Colorado, Idaho, Montana, New Mexico, Utah, Wyoming), British Columbia Brenthis myrina tollandensis Barnes & Benjamin, 1925; Ent. News 36: 44, TL: Tolland, Gilpin Co., Colorado
ab. serratimarginata (Gunder, 1926); Ent. News. 37: (1-9), TL: Vernon, British Columbia
Boloria selene tollandensis, Butterflies of America [BOA]
Brenthis myrina tollandensis ; Holland, 1928, 37; Klots, 1937, 328; McDunnough, 1938, 16
Brenthis myrina var. tollandensis ; Holland, 1931, The Butterfly Book (1931 edn): 106
Brenthis selene tollandensis ; Clark, 1941, 384
Boloria myrina tollandensis ; Brown, 1954, 64
Boloria selene tollandensis ; Remington, 1952, 67; Brown, 1957, 64; Albright, 1960, 158; Newcomer & Rogers, 1963, 171; dos Passos, 1964, 89; Shepard, 1975, in Howe, 246; Kohler, 1977, J. Lep. Soc. 31 (4): 253
Clossiana selene tollandensis ; [NACL, #4464d]

C. s. albequina (Holland, 1928)

Yukon, SW.Alaska Brenthis albequina Holland, 1928; 40, f. 4-6, TL: White Horse Pass, Yukon Territory
Brenthis albeguina var. baxteri Holland, 1928; 42
Brenthis albequina ; Holland, 1931, The Butterfly Book (1931 edn): 107, pl. LV, f. 19-21 (types); McDunnough, 1932, 269
Brenthis myrina atrocostalis ab. baxteri ; McDunnough, 1938, 16
Boloria selene albequina ab. baxteri ; dos Passos, 1964, 89
Boloria selene albequina ; dos Passos, 1964, 89; Kohler, 1977, J. Lep. Soc. 31 (4): 255, f. 4
Clossiana selene albequina ; [NACL, #4464e]

C. s. atrocostalis (Huard, 1927)

Nova Scotia, Quebec, Saskatchewan, Manitoba, Maine, Vermont, New Hampshire, New York, Michinag, Wisconsin, Minnesota, North Dakota Argynnis atrocostalis Huard, 1927; 313 (1926), TL: Chicoutini, Quebec
Brenthis myrina ab. jenningase Holland, 1928; 36
Brenthis myrina var. jenningsae Holland, 1931; The Butterfly Book (1931 edn): 105, pl. LVI, f. 9 (type)
Boloria selene atrocostalis, Butterflies of America [BOA]

Brenthis myrina atrocostalis ; McDunnough, 1930, 107; McDunnough, 1938, 16; dos Passos & Grey, 1934, 190 Brenthis myrina atrocostalis ab. jenningsae ; McDunnough, 1932, 269; McDunnough, 1938, 16 Boloria selene atrocostalis ab. jenningsae ; dos Passos, 1964, 89

Boloria selene atrocostalis ; Clark, 1941, 384; Klots, 1951, 89; dos Passos, 1954, 89; Shepard, 1975, in Howe, 246 Bororia selene atrocostalis ; Kohler, 1977, J. Lep. Soc. 31 (4): 258, f. 5, 8 Boloria selene atrocostalis ; [BNA, 318] Clossiana selene atrocostalis ; [NACL, #4464f]

C. s. terraenovae (Holland, 1928)

Newfoundland Brenthis myrina terrae-novae Holland, 1928; 36, TL: Newfoundland
Brenthis myrina terrae-novae ; dos Passos, 1935, 87; McDunnough, 1938, 16
Brenthis myrina var. terrae-novae ; Holland, 1931, The Butterfly Book (1931 edn): 105, pl. LV, f. 13 (m.type)
Brenthis selene terrae-novae ; Clark, 1941, 384
Boloria selene terrae-novae ; Klots, 1951, 89
Boloria selene terraenovae ; dos Passos, 1964, 89; Shepard, 1975, in Howe, 264; Kohler, 1977, JLS#: 362, f. 5
Clossiana selene terraenovae ; [NACL]

Clossiana perryi (Butler, 1882)

S.Ussuri, N.Korea, Amur?. See [About maps] Argynnis perryi Butler, 1882; Ann. mag. nat. Hist. (5) 9: 16
Clossiana selene perryi ; [BARU]
Clossiana perryi ; [BRU2: 46, pl. 32, f. 7-9]

Clossiana erubescens (Staudinger, 1901)

-. See [About maps] Argynnis erubescens Staudinger, 1901; in Staudinger & Rebel, Cat. Lepid. palaearct. Faunengeb., 1: 35
?Argynnis hegemone Alphéraky, 1881; Horae Soc. ent. Ross. 16 (3-4): 407, pl. 15, f. 16-17, TL: Kuldja?
Argynnis hegemone Staudinger, 1881; Stett. ent. Ztg 42: 292, TL: "Ala Tau" [Dzhungarsky Alatau Mts., Kazakhstan]
Clossiana hegemone; [BRU2: 46, pl. 33, f. 25-30]
Clossiana hegemone, NSG Voucher Specimen [Wahlberg]
Clossiana hegemone ; [BOW: pl. 194, f. 12]
Clossiana erubescens ; [Oleg Kosterin]

C. e. tianschanica (Wagner, 1913)

Tian-Shan tianschanica (Wagner, 1913)

= Clossiana hegemone hegemone (Staudinger, 1881); [BRU2, 46]

darjana (Seitz, [1909])

= Clossiana hegemone hegemone (Staudinger, 1881); [BRU2, 46]

Clossiana erubescens tienschanica ; [Oleg Kosterin]
Clossiana hegemone hegemone ; [BRU2, 46]

400x533 (~49Kb) underside An alpine meadow on the right bank terrace of the fourth tributary of the Sargir River (the Usek River basin), altitude 3200 m, 47 km NEN of Zharkent, 21 km NEN of village Sarybel', the mountain massif Tyshkantau, the Southern Dzhungarian Alatau mountain chain (NE part of Tien Shan), Zharkent District, Taldy-Kurgan Province, SE Kazakhstan. 27th of July 1994, Photo © Oleg Kosterin

C. e. houri (Wyatt, 1961)

W.Tian-Shan, Ghissar-Darvaz, Pamirs-Alai Boloria hegemone houri Wyatt, 1961; J. Lep. Soc. 15 (1): 8, TL: Bala Quran, Anjuman Valley, Hindu Kush Mts., 14300ft
Clossiana hegemone houri ; [BRU2, 46]

?C. e. chotana (O. Bang-Haas, 1915)

Argynnis erubescens chotana O. Bang-Haas, 1915; Dt. ent. Z. Iris 29: 92-100, TL: Kun-Lun Mts., China
?Clossiana hegemone chotana ; [BRU2, 46 (?)]

Clossiana selenis (Eversmann, 1837)

-. See [About maps] Argynnis selenis Eversmann, 1837; Bull. Soc. imp. Nat. Moscou 10 (1): 10, TL: "Kasan" [Kazan, Tatarstan, Russia]
?Clossiana speranda Grosser, 1979; , TL: NW.Mongolia

= ?; [BRU2, 44 (?)]

Boloria selenis, NSG Voucher Specimen [Wahlberg]
Clossiana selenis ; [BARU, #189]; [BOW: pl. 194, f. 15]; [MRS, 478]; [BRU2: 43, pl. 31, f. 25-30]
Boloria selenis ; [Otakar Kudrna]

Larva on Viola [BRU2, 44]

C. s. selenis

S.Urals, Volga Valley Clossiana selenis selenis ; [BRU2, 43]

C. s. sibirica (Erschoff, 1870)

Siberia, Amur festiva (Krulikovsky, 1893)
meinhardi (Sheljuzhko, 1929)
Clossiana selenis sibirica ; [Oleg Kosterin]; [BRU2, 44]

532x326 (~51Kb) underside A southern steppefied slope of the mountain on the Chuya River right bank between the villages Kurai and Chagan-Uzun, Altai Mts., West SIberia, Russia. 7th July 1988, Photo © Oleg Kosterin

C. s. samkoi (Sheljuzhko, 1931)

Polar Urals, Yamal Peninsula Clossiana angarensis samkoi ; [BARU]
Clossiana selenis samkoi ; [BRU2, 44]

C. s. kononovi (Kurentzov, 1970)

NE.Siberia perunovi Korb, 1999
Clossiana selenis kononovi ; [BRU2, 44]

C. s. chosensis (Matsumura, 1925)

Ussuri Clossiana selenis chosensis ; [BRU2, 44]

C. s. onorensis (Matsumura, 1925)

Sakhalin Clossiana selenis onorensis ; [BRU2, 44]

Clossiana angarensis (Erschoff, 1870)

-. See [About maps] Argynnis angarensis Erschoff, 1870; Bull. Soc. Imp. Nat. Moscou 43 (1): 112, TL: "the Irkutsk Province"
Boloria angarensis, NSG Voucher Specimen [Wahlberg]
Clossiana angarensis ; [BARU, #193]; [BOW: pl. 194, f. 10]; [BRU2: 43, pl. 30, f. 7-19]
Boloria angarensis ; [Otakar Kudrna]

533x400 (~45Kb) upperside female An open larch (Larix gmelinii (Rupr.) Rupr.)/birch (Betula pendula Roth. and B. davurica Pallas) forest above the left bluff of the Budyumkan River valley 5 km of its confluence with the Argun' River 6 km S of the village Uryupino, Gazimurskozavodskoi District, E Chita Province, E Transbaikalia, Siberia, Russia. 2nd August 1997, Photo © Oleg Kosterin

352x299 (~22Kb) underside female A road in a larch (Larix gmelinii (Rupr.) Rupr.) forest not far from the Argun' River left bank between the village Uryupino and the Gazimur River, Gazimurskozavodskoi District, E Chita Province, E Transbaikalia, Siberia, Russia. 31th July 1997, Photo © Oleg Kosterin

Larva on Vaccinium [BRU2, 43]

C. a. angarensis

Transbaikalia, S.Siberia Clossiana angarensis angarensis ; [BRU2, 43]

C. a. herzi (Wnukowsky, 1927)

N.Transbaikalia, Far East Yakutia Clossiana angarensis herzi ; [BARU]; [BRU2, 43]

C. a. sedychi (Weiss, 1964)

Polar Urals, Yamal Peninsula Clossiana angarensis sedychi ; [BRU2, 43]

C. a. alticola (Sushkin & Tschetverikov, 1907)

Sayan, Tuva (mountains) shelkovnikovi Korb, 1997
Clossiana angarensis alticola ; [BARU]; [BRU2, 43]

C. a. hakutosana (Matsumura, 1927)

Amur, Ussuri Clossiana angarensis hakutosana ; [BRU2, 43]

C. a. miakei (Matsumura, 1919)

Sakhalin Clossiana angarensis miakei ; [BARU]; [BRU2, 43]

Clossiana oscarus (Eversmann, 1844)

-. See [About maps] Argynnis oscarus Eversmann, 1844; Bull. Soc. Imp. Nat. Moscou 17 (3): 588, pl. 14, f. 1 a,b, TL: Altai [Irkutsk region?]
Boloria oscarus, NSG Voucher Specimen [Wahlberg]
Clossiana oscarus ; [BARU, #192]; [BOW: pl. 194, f. 14]; [BRU2: 44, pl. 34, f. 1-6]

488x400 (~38Kb) underside male A bushy meadow on the second right bank terrace of the Koen River at the Volchikha Rivulet just upstream of its fall into the Koen River near the village Nizhniy Koen, Iskitim District, Novosibirsk Province, West Siberia, Russia. 18th June, 1989, Photo © Oleg Kosterin

Larva on Viola [BARU]

C. o. oscarus

S.Siberia, E.Amur Clossiana oscarus oscarus ; [BRU2, 44]

C. o. australis (Graeser, 1888)

S.Amur, Ussuri, Sakhalin maxima (Fixsen, 1887); (preocc.)
major (Graeser, 1888)
Clossiana oscarus australis ; [BARU]; [BRU2, 44]

C. o. oscaroides (Ménétriés, 1859)

E.Yakutia Clossiana oscarus oscaroides ; [BRU2, 44]

Clossiana freija (Thunberg, 1791)

Japan. See [About maps] Muurainhopeatäplä Freija's Fritillary Freya's Fritillary Zigzag Fritillary Frejas pärlemorfjäril Myrpärlefjäril
Papilio freija Thunberg, 1791; Diss. Ent. sistens Insecta Suecica (1) (2): 34, TL: Vestrobothnia & Lapponica
?Papilio freija Beclin, 1791
freya (Godart, 1819)
gunderi (Harper, 1933)
Boloria freija freija, Butterflies of America [BOA]
Boloria freija, NSG Voucher Specimen [Wahlberg]
Boloria freija, Butterflies and Moths of North America [BAMONA]
Boloria freija, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Boloria freija, Svenska fjärilar [SVF]
Boloria freija, Norges Sommerfugler [NORLEP]
Clossiana freija, Moths and Butterflies of Europe [MBE]
Brenthis freija ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 16
Clossiana freija ; [BARU, #201]; [NACL, #4471]; [MRS, 477]; [BRU2: 48, pl. 32, f. 24-29]
Boloria freija ; [BNA: 323, #249, pl. 26]; [BOC, 284]; [Opler]; [Otakar Kudrna]

Larva on Rubus chamaemorus, Vaccinium uliginosum [SPRK], Vaccinium vitis-idaeus, Rhododendron auerum [BARU], Empetrum nigrum [MRS], Vaccinium caespitosum, V. uliginosum, Arctostaphylos uva-ursi, Empetrum nigrum [NAB], Arctous alpina, (Scandinavia) Sieversia, Rhododendron, (Japan) Dryas integrifolia, (N.America) [BRU2, 48]

C. f. freija

Fennoscandia, W.Siberia, C.Siberia, Far East, Transbaikiali, Chukotka, Amur, Ussuri, Canada, Alaska Boloria freija freija
freija (Quensel, 1791); (preocc.)
lapponica (Esper, 1793)
Boloria freija freija ; [BNA, #249]
Clossiana freija freija ; [BRU2, 48]

C. f. pallida (Elwes, 1899)

Altai, Sayan, Tuva calais (Fruhstorfer, 1916)
Clossiana freija pallida ; [BARU]
Clossiana freija calais ; [BARU, (Tuva)]
Clossiana freija pallida ; [Oleg Kosterin]; [BRU2, 48]

533x400 (~67Kb) underside female A dwarf birch (Betula rotundifolia Spach) montane tundra at 2200 m above sea level in the Argem (Direntai) Stream basin, 15 km W of the village Dzhazatr, on the eastern spurs of the Katunskiy Mt. Range, Kosh-Agach District, Central Altai Mts., West Siberia, Russia. 13th July, 1988, Photo © Oleg Kosterin

508x444 (~37Kb) upperside male A meadowy-tundrous SW slope of a ridge between the headwaters of the Chikty Rivulet, 2600 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 10th July 1998 (on Aster alpinum L.), Photo © Oleg Kosterin

C. f. jakutensis (Wnukowsky, 1927)

NE.Yakutia Clossiana freija jakutensis ; [BARU]; [BRU2, 48]

C. f. tarquinius (Curtis, 1835)

freija tarquinius (Curtis, 1835); in Ross, Nar. 2nd Voy. N.-W. Pass., App.: 68
Brenthis freija tarquinius ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 16
Clossiana freija tarquinius ; [NACL, #4471a]

C. f. browni (Higgins, 1953)

Rocky Mountains Boloria freija browni, Butterflies of America [BOA]
Clossiana freija browni ; [NACL, #4471d]
Boloria freija browni ; [BNA, #249]

Clossiana natazhati (Gibson, 1920)

Yukon, Victoria I.. See [About maps] Pleistocene Fritillary
Brenthis natazhati Gibson, 1920; Rep. Canadian arctic expedition 1913-18. 3 (1): 21i-22i, TL: Mt. Natazhat, St. Elias Mts., Yukon Territory, 2470-2620m
Boloria freija nabokovi Stallings & Turner, 1947; Can. Entomol. 78 (7-8): (134-135), TL: Alaska Military Highway, 102 miles N of Summit 2, 1830m

= ; Troubridge & Wood, 1990, J. Lep. Soc. 44 (3): 181

Boloria natazhati natazhati, Butterflies of America [BOA]
Boloria natazhati, Butterflies and Moths of North America [BAMONA]
Boloria freija nabokovi ; dos Passos, 1964; Howe, 1975
Clossiana freija nabokovi ; Miller & Brown, 1981; [NACL, #4471c]; Tilden & Smith, 1986
Boloria freija natazhati ; dos Passos, 1964; Howe, 1975
Clossiana freija natazhati ; Miller & Brown, 1981; Ferris, dos Passos, Ebner & Lafontaine, 1983; [NACL, #4471b]; Tilden & Smith, 1968
Boloria natazhati ; [BNA: 323, #250, pl. 24]; Troubridge & Wood, 1990, J. Lep. Soc. 44 (3): 181, f. 1-2; [Opler]

Larva on Dryas integrifolia? Troubridge & Wood, 1990, J. Lep. Soc. 44 (3): 185

Clossiana polaris (Boisduval, [1828])

Arctic Europe, Arctic Asia, Arctic America. See [About maps] Pohjanhopeatäplä Polar Fritillary Polaris Fritillary Polarpärlefjäril
polaris (Boisduval, [1828]); Eur. Lepid. Index Meth. 1: 15, TL: Nordkap Peninsula [Norway]
groenlandica (Skinner, 1892)
americana (Strand, 1905)
Boloria polaris polaris, Butterflies of America [BOA]
Boloria polaris, NSG Voucher Specimen [Wahlberg]
Boloria polaris, Butterflies and Moths of North America [BAMONA]
Boloria polaris, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Boloria polaris, Svenska fjärilar [SVF]
Brenthis polaris ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 16
Clossiana polaris ; [NACL, #4470]; [BOW: pl. 2, f. 1]; [BARU, #202]; [BRU2: 48, pl. 33, f. 1-3]
Boloria polaris ; [BNA: 323, #248, pl. 26]; [Opler]; [Otakar Kudrna]

660x498 (~61Kb) upperside female FINLAND: Li: Utsjoki Puollamoaivi, 775:50, 8.7.1997, Photo © Markku Savela

Larva on Cassiope tetragona, Vaccinium uliginosum, Dryas octopetala, D. punctata [BRU2, 48]

C. p. stellata (Masters, 1972)

Clossiana polaris stellata ; [NACL, #4470a]

Clossiana thore (Hübner, [1803-1804])

Japan. See [About maps] Purohopeatäplä Thore's Fritillary Gråkantad pärlemorfjäril Bäckpärlefjäril Alpen-Perlmutterfalter
Papilio thore Hübner, [1803-1804]; Samml. eur. Schmett. [1] : pl. 111 (1803-1804), f. 571-573, TL: Alps of Tirol
Boloria thore, NSG Voucher Specimen [Wahlberg]
Boloria thore, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Boloria thore, Svenska fjärilar [SVF]
Clossiana thore, Moths and Butterflies of Europe [MBE]
Clossiana thore, Europaische Schmetterlinge [Mario Maier]
Clossiana thore ; [BOW: pl. 2, f. 4]; [BARU, #194]; [MRS, 478]
Boloria thore ; [BRU2: 46, pl. 29, f. 22-24]; [Otakar Kudrna]

374x506 (~29Kb) underside Among the dead branches of the Siberian dwarf pine (Pinus pumila Ledeb.) A forest/tundra on the Koni Peninsula, S Magadan Region, Russia. 11th of July 1989, Photo © Oleg Kosterin

494x684 (~70Kb) upperside

470x733 (~69Kb) underside

Larva on Viola bifolia, (Finland) Viola altaica, (Urals) [BRU2, 47] Viola selkirkii, Viola grypoceras [MRS]

C. t. thore

Alps Clossiana thore thore ; [BRU2, 46]

C. t. exellens (Krulikovsky, 1893)

splendida (Krulikovsky, 1895)
Clossiana thore exellens ; [BRU2, 47]

C. t. borealis (Staudinger, 1861)

N.Scandinavia, N.Siberia, Far East, Kamchatka Clossiana thore borealis ; [BRU2, 47]

C. t. transuralensis (Sheljuzhko, 1931)

Ural, W.Siberia Clossiana thore transuralensis ; [BARU]; [BRU2, 47]

C. t. hypercala (Fruhstorfer, 1907)

Altai, Sayan, Transbaikalia thore hypercala (Fruhstorfer, 1907) (Argynnis); Ent. Zs. 21 (27): 163
lacrimosa (Fruhstorfer, 1907)
Clossiana thore hypercata ; (spell.?)

C. t. herculea (Fruhstorfer, 1907)

? herculea Fruhstorfer, 1907; , TL: Altai
Clossiana thore herculea ; [BARU]

C. t. hyperusia (Fruhstorfer, 1907)

Amur, Ussuri thore hyperusia (Fruhstorfer, 1907) (Argynnis); Ent. Zs. 21 (27): 163
meridionalis (Kardakov, 1928)

= ; [BRU2, 47]

Clossiana brenda Hemming, 1933

= ; [BRU2, 47]

Clossiana thore meridionalis ; [BARU]
Clossiana thore hyperusia ; [BARU]; [MRS, 478]; [BRU2, 47]

C. t. karafutonis (Matsumura, 1919)

Sakhalin Clossiana thore karafutonis ; [BRU2, 47]

Clossiana hakutozana (Matsumura, 1927)

N.Korea, NE.China. See [About maps] hakutozana (Matsumura, 1927); , TL: N.Korea
Clossiana hakutozana ; [BARU, #195]
Clossiana hakutosana ; [EBW, (name only)]

Clossiana frigga (Thunberg, 1791)

. See [About maps] Rahkahopeatäplä Frigga's Fritillary Willow-Bog Fritillary Friggas pärlemorfjäril Hjortronpärlefjäril
Papilio frigga Thunberg, 1791; Diss. Ent. sistens Insecta Suecica (1) (2): 33, TL: Lappland
?Papilio frigga Becklin, 1791
Boloria frigga, NSG Voucher Specimen [Wahlberg]
Boloria frigga, Butterflies and Moths of North America [BAMONA]
Boloria frigga, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Boloria frigga, Svenska fjärilar [SVF]
Clossiana frigga, Moths and Butterflies of Europe [MBE]
Brenthis frigga ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 16
Clossiana frigga ; [NACL, #4466]; [BARU, #196]; [BOW: pl. 1, f. 27]; [BRU2: 47, pl. 32, f. 13-23]
Boloria frigga ; [BNA: 321, #244, pl. 26]; [Opler]; [Otakar Kudrna]

461x381 (~37Kb) upperside

447x373 (~36Kb) underside

Larva on Rubus chamaemorus [SPRK], Salix, Betula, Dryas integrifolia [BNA, 321]

C. f. frigga

N.Scandinavia helvenacius (Sedykh, 1977)
Clossiana frigga frigga ; [BRU2, 47]

C. f. alpestris (Elwes, 1899)

Altai, Sayan, Transbaikalia Clossiana frigga alpestris ; [BARU]; [BRU2, 47]

872x600 (~56Kb) upperside female Highland dwarf birch tundras on of the southerm principal slope of the Yuzhno-Chuiskii mountain at the the Chikty Rivulet headwaters, 2400 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 15th July 1998, Photo © Oleg Kosterin

900x592 (~98Kb) upperside female A flowery alpine meadow in a valley of the rivulet descending from the southerm principle slope of the Yuzhno-Chuiskii mountain range between the Chikty and Akbul Rivulets, 2600 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 11th July 1998, Photo © Oleg Kosterin

475x500 (~33Kb) underside male An alpine meadow vegetation at a brook flowing in a moss/sedge tundra in a cirque of the sources of a rivulet descending from the southerm principle slope of the Yuzhno-Chuiskii mountain range between the Chikty and Akbul Rivulets, 2650 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 16 July 1998, Photo © Oleg Kosterin

C. f. maritima (Kardakov, 1928)

Amur, Ussuri Clossiana frigga maritima ; [BARU]; [BRU2, 47]

C. f. annae (Sushkin, 1906)

Tarbgatai Mts. Clossiana frigga annae ; [BRU2, 47 (?)]

C. f. saga (Staudinger, 1861)

Canada ?saga Kaden, 1861; Stett. Ent. Ztg. ?23: 350
Brenthis frigga saga Kaden, 1861
Brenthis frigga saga ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 16
Clossiana frigga saga ; [NACL, #4466a]
Boloria frigga saga ; [BNA, #244]

C. f. gibsoni (Barnes & Benjamin, 1926)

Alaska - Hudson Bay alaskensis (Lehmann, 1913)
gibsoni (Barnes & Benjamin, 1926)
lehmanni (Holland, 1928)
Boloria frigga gibsoni, Butterflies of America [BOA]
Clossiana frigga alaskensis ; [NACL, #4466b]
Boloria frigga gibsoni ; [BNA, #244]

C. f. sagata (Barnes & Benjamin, 1923)

Wyoming - Colorado Boloria frigga sagata, Butterflies of America [BOA]
Clossiana frigga sagata ; [NACL, #4466c]
Boloria frigga sagata ; [BNA, #244]

Clossiana dia (Linnaeus, 1767)

-. See [About maps] Violet Fritillary Hainveilchen-Scheckling La Petite Violette Perlada Violeta Dostojka dia
Papilio dia Linnaeus, 1767; Syst. Nat. (Edn 12) 1 (2): 785, TL: Austria
Clossiana dia, NSG Voucher Specimen [Wahlberg]
Boloria dia, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Clossiana dia, Moths and Butterflies of Europe [MBE]
Clossiana dia, Europaische Schmetterlinge [Mario Maier]
Clossiana dia, Captain's European Butterfly Guide [Simon Coombes]
Clossiana dia ; [BOW: pl. 1, f. 28]; [EBW]; [BARU, #200]; [MRS, 477]; [BRU2: 49, pl. 33, f. 13-21]
Boloria dia ; [Otakar Kudrna]

533x400 (~44Kb) underside A dwarf birch (Betula rotundifolia Spach) montane tundra at 2200 m above sea level in the Argem (Direntai) Stream basin, 15 km W of the village Dzhazatr, on the eastern spurs of the Katunskiy Mt. Range, Kosh-Agach District, Central Altai Mts., West Siberia, Russia. 129 - 15th, 130 - 12th July, 1988, Photo © Oleg Kosterin	598x400 (~44Kb) larva Central Europe 1990-2000, Photo © Michael Borsch	598x400 (~45Kb) upperside Central Europe 1990-2000, Photo © Michael Borsch
465x500 (~43Kb) upperside Axat, Pyrenees orientales, France 24/07/96, Photo © Simon Coombes	500x503 (~50Kb) underside Axat, Pyrenees orientales, France 24/07/96, Photo © Simon Coombes

Larva on Viola, Rubus idaeus, Prunella vulgaris [BRU2, 49]

C. d. dia

W.Europe Clossiana dia dia ; [BRU2, 49]

C. d. disconota (Krulikovsky, 1909)

CEU, W.Siberia Clossiana dia disconota ; [BRU2, 49]

C. d. calida (Jachontov, 1911)

Caucasus, Transcaucasus Clossiana dia calida ; [BRU2, 49]

C. d. alpina (Elwes, 1899)

Dzhungarsky Alatau, Altai, Sayan, Transbaikalia dia alpina (Elwes, 1899); , TL: Altai
Clossiana dia alpina ; [BARU]; [BRU2, 49]

C. d. semota Tuzov, 2000

Far East Clossiana dia semota ; [BRU2, 49]

C. d. setania (Fruhstorfer, 1909)

dia setania (Fruhstorfer, 1909) (Argynnis); Int. ent. Zs. 3 (4): 21, TL: 'Marokko' [mislabeling?]
Clossiana dia setania ; [BMAT: 105, pl. 32, f. 19-20 (not in N.Africa)]

Clossiana gong (Oberthür, 1884)

Tibet, W.China, N.China. See [About maps] ?gong (Oberthür, 1914); Etudes d'Ent. 9: 15
Clossiana gong ; [BOW: pl. 194, f. 11]; [MRS, 477]

C. g. splendens Murayama

Clossiana gong splendens ; [MRS, 477]

C. g. xizangensis Huang, 2000

Clossiana jerdoni (Lang, 1868)

W.Himalayas, Chitral - Kashmir. See [About maps] Jerdons Silverspot
Argynnis jerdoni Lang, 1868; Ent. Montrhly Mag. 5: 34
Clossiana jerdoni ; [BOW: pl. 194, f. 13]

Clossiana improba (Butler, 1877)

. See [About maps] Kääpiöhopeatäplä Dusky-windged Fritillary Dinky Fritillary Dingy Arctic Fritillary Dvärgpärlefjäril
Argynnis improba Butler, 1877; Ent. mon. Mag. 13: 206, TL: "Winter Cove and Cambridge Bay" [NW Territories, Canada]
Boloria improba, Butterflies and Moths of North America [BAMONA]
Boloria improba, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Boloria improba, Svenska fjärilar [SVF]
Clossiana improba, Moths and Butterflies of Europe [MBE]
Brenthis frigga improba ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 16
Clossiana improba ; [NACL, #4467]; [BARU, #197]; [BOW: pl. 13, f. 20]; [BRU2: 47, pl. 33, f. 22-24]
Boloria improba ; [BNA: 321, #245, pl. 5, 26, 60]; [Opler]; [Otakar Kudrna]

600x450 (~38Kb) Gohpascurro/Bjorkliden, 1100-1400m, Torne Lapmark, Sweden 9 June 1985, Photo © Lars Andersen	800x583 (~75Kb) underside Gohpascurro/Bjorkliden, 1100-1400m, Torne Lapmark, Sweden 9 June 1985, Photo © Lars Andersen	800x549 (~56Kb) upperside Gohpascurro/Bjorkliden, 1100-1400m, Torne Lapmark, Sweden 9 June 1985, Photo © Lars Andersen
800x709 (~76Kb) upperside Gohpascurro/Bjorkliden, 1100-1400m, Torne Lapmark, Sweden 9 June 1985, Photo © Lars Andersen

Larva on Bistorta vivipara, Salix herbacea [BRU2, 47], Polygonum viviparum, Salix [BARU], Salix, S. reticulata?, (NW.Terr.) [BNA, 321]

C. i. improba

Arctic Europe, Arctic Asia Clossiana improba improba ; [BRU2, 47]

C. i. improbula (Bryk, 1920)

C. i. youngi (Holland, 1900)

Alaska, E.Chukotka? improba youngi (Holland, 1900); Ent. News 11: 283, TL: Alaska
Brenthis youngi ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 16
Clossiana improba youngi ; [NACL, #4467a]

C. i. harryi

Wyoming (Wind River Mts.) Boloria improba harryi
Boloria improba harryi, Butterflies of America [BOA]
Boloria improba harryi ; [BNA, #245]

Larva on Salix arctica [BNA, 321]

?C. i. acrocnema (Gall & Sperling, 1980)

Colorado (San Juan Mts.) Uncompaghre Fritillary
Boloria acrocnema Gall & Sperling, 1980; J. Lep. Soc. 43: (230-252), TL: near Uncompahgre Peak, San Juan Mts., Hinsdale Co., Colorado
Boloria improba acrocnema, Butterflies of America [BOA]
Boloria improba acrocnema ; [BNA, #245]

Larva on Salix nivalis [BNA]

Clossiana titania (Esper, 1793)

. See [About maps] Lehtohopeatäplä Titania's Fritillary Purple Bog Fritillary Amathusias pärlemofjäril Lundpärlefjäril Natterwurz-Perlmutterfalter Dostojka titania
Papilio titania Esper, 1793; Die Schmett. 1 (Suppl.): 58, TL: Sardinia
Boloria titania, NSG Voucher Specimen [Wahlberg]
Boloria titania, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Clossiana titania, Moths and Butterflies of Europe [MBE]
Clossiana titania, Europaische Schmetterlinge [Mario Maier]
Clossiana titania, Captain's European Butterfly Guide [Simon Coombes]
Clossiana titania ; [BOW: pl. 2, f. 5]; [EBW]; [BARU, #198]; [BRU2: 49, pl. 33, f. 31-36]
Boloria titania ; [Otakar Kudrna]

Misidentified/misapplied:

Clossiana titania (= Clossiana chariclea); [NACL, #4474]
Boloria titania (= Clossiana chariclea); [BNA: 319, #240]

C. t. titania

Sardinia, ?

C. t. bivina (Fruhstorfer, 1908)

CEU, SEU titania bivina (Fruhstorfer, 1908) (Argynnis); Int. ent. Zs. 1 (41): 310
amathusia (Esper, [1784]); (preocc.)
rossica (Hemming, 1934)
Clossiana titania bivina ; [BARU]; [BRU2, 50]

Larva on Viola, Vaccinium uliginosum [BRU2, 50]

C. t. staudingeri (Wnukowsky, 1929)

S.Siberia, Altai, Sayan, mountains (Tuva), Transbaikalia, Amur, Sakhalin? sibirica (Staudinger, 1892); (preocc.)
altaica (Seitz, [1909]); (preocc.)
julia (Hemming, 1934)
Clossiana titania staudingeri ; [BARU]; [BRU2, 50]

Larva on Bistorta major, Filipendula ulmaria, Trollius asiaticus [BRU2, 50]

C. t. miyakei (Matsumura, 1919)

Sakhalin, Low Amur Clossiana titania miyakei ; [BARU]

Clossiana euphrosyne (Linnaeus, 1758)

. See [About maps] Pursuhopeatälä Pearl-bordered Fritillary Prydlig pärlemorfjäril Skogspärlefjäril Silberstrich Grand Collier Argenté Perlada rojiza Dostojka eufrozyna
Papilio euphrosyne Linnaeus, 1758; Syst. Nat. (Edn 10) 1: 481, no. 142
Papilio euphrosyne Linnaeus, 1761; Fauna Suecica (Edn 2): 282, TL: Sweden
Clossiana euphrosyne, NSG Voucher Specimen [Wahlberg]
Boloria euphrosyne, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Boloria euphrosyne, Svenska fjärilar [SVF]
Boloria euphrosyne, Norges Sommerfugler [NORLEP]
Clossiana euphrosyne, Moths and Butterflies of Europe [MBE]
Clossiana euphrosyne, Europaische Schmetterlinge [Mario Maier]
Clossiana euphrosyne, Captain's European Butterfly Guide [Simon Coombes]
Clossiana euphrosyne ; [BOW: pl. 1, f. 26]; [EBW]; [BARU, #190]; [BRU2: 44, pl. 31, f. 13-24]
Boloria euphrosyne ; [Otakar Kudrna]

500x349 (~71Kb) upperside Luckett, Cornwall, UK 20/05/95, Photo © Simon Coombes	417x500 (~57Kb) underside Luckett, Cornwall, UK 20/05/95, Photo © Simon Coombes	533x400 (~50Kb) larva An open pine grove above the bluff of the valley of the Bol'shoi Elbash River, 5 km NW of the village Ust'-Chem, Iskitim dist., Novosibirsk region, West Siberia, 9th May 1995 (mature larva), Photo © Oleg Kosterin
600x832 (~62Kb) underside Finland, Loppi, Luutasuo 673:35, 2.6.1979, Photo © Pertti Pakkanen	900x745 (~146Kb) upperside female FINLAND: Li: Inari, Luttojoki, 760:54, 11.7.1998, Photo © Markku Savela	600x722 (~61Kb) upperside FINLAND, Ab: Suomusjärvi, 669:31, 15.6.1997, Photo © Markku Savela
900x1291 (~125Kb) Finland, Ab: Suomusjärvi (669:31), 6.6.2004, Photo © Markku Savela	800x631 (~57Kb) underside FINLAND: Ka: Virolahti, 671:53 (Reskopinsuo), 14.6.1997, Photo © Markku Savela

Larva on Viola, V. palustris, V. riviniana, Vaccinium uliginosum, Ledum palustre [SPRK], Viola hirta, Waldsteinia ternata [BRU2, 45]

C. e. euphrosyne

CEU, Siberia Clossiana euphrosyne euphrosyne ; [BRU2, 45]

C. e. fingal (Herbst, 1804)

NEU, Siberia Clossiana euphrosyne fingal ; [BRU2, 45]

C. e. rusalka (Fruhstorfer, 1909)

SEU, W.Siberia euphrosyne rusalka (Fruhstorfer, 1909) (Argynnis); Int. ent. Zs. 3 (20): 112
Clossiana euphrosyne rusalka ; [BRU2, 45]

C. e. orphana (Fruhstorfer, 1907)

Transbaikalia, Amur, Ussuri Argynnis euprhosyne orphanus Fruhstorfer, 1907; Ent. Zs. 21 (27): 164
Clossiana euphrosyne orphanus ; [BARU]
Clossiana euphrosyne orphana ; [BRU2, 45]

C. e. kamtschadalus (Seitz, [1909])

Kamchatka, N.Sakhalin euphrosyne kamtschadalus (Seitz, [1909]); , TL: Kamchatka
Clossiana euphrosyne kamtschadalus ; [BARU]; [BRU2, 45]

C. e. umbra (Seitz, [1909])

Altai, Sayan Clossiana euphrosyne umbra ; [BRU2, 45]

C. e. dagestanica (Sovinsky, 1905)

Caucasus, Transcaucasia euphrosyne dagestanica (Sovinsky, 1905) (Argynnis); Russk. ent. Obozr. 5 (3-4): 108
anka (Fruhstorfer, 1907) (Argynnis); Ent. Zs. 21 (27): 164
Clossiana phaenea Jachontov, 1908

?C. e. nephele (Herrich-Schäffer, [1847])

Ural, Siberia euphrosyne nephele (Herrich-Schäffer, [1847]) (Argynnis); Syst. Bearb. Schmett. Europ. 1 (26): (ii) f. 371-372
Clossiana euphrosyne nephele ; [BARU]

Clossiana chariclea (Schneider, 1794)

. See [About maps] Tundrahopeatäplä Arctic Fritillary Tundrapärlefjäril
chariclea (Schneider, 1794); Neuest. Mag. Lieb. Ent. 1 (5): 588, TL: Lappland
Clossiana chariclea, Lepidoptera collection [SZMN]
Boloria chariclea, NSG Voucher Specimen [Wahlberg]
Boloria chariclea, Butterflies and Moths of North America [BAMONA]
Boloria chariclea, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Boloria chariclea, Svenska fjärilar [SVF]
Clossiana carichlea ; Zetterstedt, 1839, (missp.)
Clossiana churiclea ; Johansen, 1921, (missp.)
Clossiana churchilea ; dos Passos, 1964, (missp.)
Brenthis chariclea ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 16
Clossiana chariclea ; [NACL, #4475]; [BARU, #199]; [BOW: pl. 1, f. 25]; [BRU2: 48, pl. 33, f. 4-9]
Boloria chariclea ; [BOC, 286]; [Opler]; [Otakar Kudrna]

550x424 (~48Kb) upperside FINLAND: Li: Utsjoki Puollamoaivi, 775:50, 8.7.1997, Photo © Markku Savela

500x564 (~35Kb) underside FINLAND: Li: Utsjoki Ailigas, 775:50, 8.7.1997, Photo © Markku Savela

700x506 (~69Kb) upperside FINLAND: Li: Utsjoki Ailigas, 775:50, 6.7.1997, Photo © Markku Savela

Larva on Dryas integrifolia, Salix arctica, S. reticulata, (N.America) [BRU2, 49]

C. c. chariclea

Arctic Europe Clossiana chariclea chariclea ; [BRU2, 48]

Larva on Cassiope tetragona [BRU2, 49]

C. c. arctica (Zetterstedt, 1839)

Arctic Asia, Wrangel Is., Chukotka chariclea arctica (Zetterstedt, 1839); Insecta Lapponica : 899
Cclossiana chariclea tshuktsha Korshunov, 1998
oenome (Scudder, 1875)
obscurata (McLachlan, 1878); J. Linn. Soc. Lond. 14: 110
Brenthis chariclea arctica ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 16
Brenthis chariclea obscurata ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 16
Clossiana chariclea arctica ; [NACL, #4475a]

C. c. butleri (Edwards, 1883)

Arctic America, Chukotka, Kamchatka? chariclea butleri (Edwards, 1883); , TL: "Cape Thompson, Kotzebue Sound, W.Alaska"
Clossiana chariclea butleri ; [BARU]; [NACL, #4475b]
Clossiana butlerii ; Edwards, 1884, (missp.)
Clossiana butleri ; [BRU2: 49, pl. 33, f. 10-12]

C. c. boisduvalii (Duponchel, 1832)

Alaska, Alberta, Labrador, Newfoundland, Minnesota, British Columbia chariclea boisduvalii (Duponchel, 1832); in Godart, Hist. nat. Lépid. Fr., Suppl. 1 (Diurnes): pl. 20, f. 4
labradorensis (Gunder, 1928)
Brenthis chariclea boisduvalii ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 16
Clossiana titania boisduvalii ; [NACL, #4474a]
Boloris titania boisduvalii ; [BNA, #240]

C. c. ranieri (Barnes & McDunnough, 1913)

Washington Brenthis chariclea ranieri Barnes & McDunnough, 1913; Contr. nat. Hist. Lep. N. Am. 2 (3): pl. II, f. 1-4, TL: Paradise Valley, Mt. Ra[i]nier, 6-7000ft, [Pierce or Lewis Co.] Wash[ington]
Boloria chariclea rainieri, Butterflies of America [BOA]
Brenthis chariclea rainieri ; Barnes & McDunnough, 1913, Contr. nat. Hist. Lep. N. Am. 2 (3): (orig. missp.)
ranieri ; Guppy & Kondla, 2002, J. Lep. Soc. 56 (3): 180
Clossiana titania rainieri ; [NACL, #4474b (missp.)]
Boloria chariclea rainieri ; [BOC, 286]

C. c. grandis (Barnes & McDunnough, 1916)

N.British Columbia, Ontario chariclea grandis (Barnes & McDunnough, 1916); , TL: Hymers, Ontario
Boloria chariclea grandis, Butterflies of America [BOA]
Clossiana titania grandis ; [NACL, #4474c]
Boloria chariclea grandis ; [BOC, 286]

C. c. montina (Scudder, 1863)

montinus (Scudder, 1863); Proc. Essex Inst. 3: 166
Brenthis montinus ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 16; dos Passos, 1936, Can. Ent. 68: 239-241
Clossiana titania montina ; [NACL, #4474d]

C. c. ingens (Barnes & McDunnough, 1918)

martini (Gunder, 1934)
Clossiana titania ingens ; [NACL, #4474e]

C. c. helena (Edwards, 1871)

Rocky Mountains chariclea helena (Edwards, 1871); Trans. amer. ent. Soc. 3: 268, TL: Mosquito Pass, Park/Lake cos, CO
Boloria chariclea helena, Butterflies of America [BOA]
Brenthis helena ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 16
Clossiana titania helena ; [NACL, #4474f]
Boloria titania helena ; [BNA, #240]
Boloria chariclea helena ; [BOC, 286]

Clossiana bellona (Fabricius, 1775)

Rocky Mountains, Canada, Washington, Oregon. See [About maps] Meadow Fritillary
Papilio bellona Fabricius, 1775; Syst. Ent.: 517
fasciata (Cockerell, 1889)
kleeni (Watson, 1921)
ammiralis (Hemming, 1933)
pardopsis (Holland, 1928)
Boloria bellona bellona, Butterflies of America [BOA]
Boloria bellona, NSG Voucher Specimen [Wahlberg]
Boloria bellona, Butterflies and Moths of North America [BAMONA]
myrina ; Martyn, 1797
Brenthis bellona ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 16
Clossiana bellona ; [NACL, #4465]; [BOW: pl. 13, f. 19]
Boloria bellona ; [BNA: 320, #243, pl. 26]; [BOC, 282]; [Opler]

Larva on Viola sororia, V. pallens [BNA, 320]

C. b. bellona

Boloria bellona bellona
Boloria bellona bellona ; [BNA, #243]

C. b. toddi (Holland, 1928)

Labrador, N.Manitoba, Northwest Territories bellona toddi (Holland, 1928); , TL: St. Margaret's River, Quebec
Clossiana bellona toddi ; [NACL, #4465a]
Boloria belona toddi ; [BNA, #243]

C. b. jenistae (D. Stallings & Turner, 1947)

Clossiana bellona jenistae ; [NACL, #4465b]

Clossiana kriemhild (Strecker, 1879)

Montana, Wyoming. See [About maps] Relict Fritillary Kriemhild Fritillary
kriemhild (Strecker, 1879); in Ruffner, Rept. Surv. Missouri. app.: ?
laurenti (Skinner, 1913)
Clossiana luecki Reuss, 1923
Boloria kriemhild, Butterflies of America [BOA]
Boloria kriemhild, Butterflies and Moths of North America [BAMONA]
Clossiana kriemeld ; McHenry, 1964, (missp.)
Clossiana lucki ; dos Passos, 1964, (missp.)
Brenthis epithore kreimhild[sic] ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 16
Clossiana kriemhild ; [NACL, #4468]
Boloria kriemhild ; [BNA: 320, #241, pl. 26]; [Opler]

Larva on Viola [NAB]

Clossiana epithore (Edwards, 1864)

Yukon - California. See [About maps] Western Meadow Fritillary Pacific Fritillary
epithore (Edwards, 1864); Proc. ent. Soc. Philad. 2: 504, TL: Saratoga, Santa Clara, California
?epithore Boisduval, 1869; Ann. Soc. ent. Belg. 12: 58
Brenthis epithore Boisduval, 1869
Boloria epithore, Butterflies of America [BOA]
Boloria epithore, Butterflies and Moths of North America [BAMONA]
Brenthis epithore ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 16
Clossiana epithore ; [NACL, #4469]
Boloria epithore ; [BNA: 320, #242, pl. 26]; [BOC, 283]; [Opler]

identification uncertain 600x539 (~44Kb) upperside USA: Near Crystal Mtn, (46°57'10N 121°28'27W), Pierce Co., WA, 27.7.1999, Photo © Markku Savela

identification uncertain 400x369 (~20Kb) underside USA: Near Crystal Mtn, (46°57'10N 121°28'27W), Pierce Co., WA, 27.7.1999, Photo © Markku Savela

Larva on Viola sempervirens, V. glabella, V. ocellata, V. adunca, V. nephrophylla? [BOC, 283]

C. e. epithore

California Boloria epithore epithore
Boloria epithore epithore ; [BNA, #242]

C. e. chermocki (E. & S. Perkins, 1966)

Oregon epithore chermocki (E. & S. Perkins, 1966); , TL: 2.9 mi E of Dolph, Yamhill Co., Oregon
Boloria epithore chermocki, Butterflies of America [BOA]
Clossiana epithore chermocki ; [NACL, #4469a]
Boloria epithore chermocki ; [BNA, #242]

C. e. uslui (Koçak, 1984)

British Columbia Boloria epithore uslui Koçak, 1984; (preocc.)
epithore borealis (E. Perkins, 1973); (preocc.), TL: Shingle Creek Rd., Keremeos, BC
obscuripennis (Gunder, 1926); (ab.)
Clossiana epithore borealis ; [NACL, #4469b]

C. e. sierra (E. Perkins, 1973)

California epithore sierra (E. Perkins, 1973); , TL: Sentinel Dome, Yosemite Nat. Park, California
eldorado (Strand, 1915); (ab.)
wawnae (Gunder, 1924); (ab.)
Clossiana epithore sierra ; [NACL, #4469c]

Clossiana alberta (Edwards, 1890)

-. See [About maps] Alberta Fritillary Alberta Alpine Fritillary
Argynnis alberta Edwards, 1890; Can. Ent. 22: 113, TL: Laggan, Alberta, Canada
banffensis (Gunder, 1932)
Boloria alberta, Butterflies of America [BOA]
Boloria alberta, Butterflies and Moths of North America [BAMONA]
Brenthis alberta ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 16
Clossiana alberta ; [NACL, #4472]; [BARU, #204]; [BOW: pl. 13, f. 17]; [BRU2: 50, pl. 35, f. 1-3]
Boloria alberta ; [BNA: 322, #246, pl. 26]; [Opler]

1600x1200 (~262Kb) upperside female Canada, Mt. Tripoli, 18.7.1989, Photo © Norbert Kondla leg.	1600x1200 (~248Kb) underside female Canada, Mt. Tripoli, 18.7.1989, Photo © Norbert Kondla leg.	1600x1200 (~233Kb) upperside male Canada, Mt. Tripoli, 18.7.1989, Photo © Norbert Kondla leg.
1600x1200 (~226Kb) underside male Canada, Mt. Tripoli, 18.7.1989, Photo © Norbert Kondla leg.

Larva on Dryas octopetala, Vaccinium [BARU], Dryas octopetala, Dryas integrifolia [BRU2, 50]

C. a. alberta

Alberta, British Columbia

C. a. kurentzovi (Wyatt, 1961)

Chukotka alberta kurentzovi (Wyatt, 1961); , TL: Chukotka
Clossiana alberta kurentzovi ; [BARU]; [BRU2, 50]

Clossiana astarte (Doubleday, [1848])

Alberta, British Columbia - Alaska, Washington, Montana. See [About maps] Astarte Fritillary Arctic Ridge Fritillary
astarte (Doubleday, [1848]); Gen. diurn. Lep. (1): pl. 53, f. 5, TL: Rock Lake, Alberta
Argynnis victoria Edwards, 1891; Can. Ent. 23: (195-199)
Boloria astarte astarte, Butterflies of America [BOA]
Boloria astarte, Butterflies and Moths of North America [BAMONA]
Brenthis astarte ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 16
Clossiana astarte ; [NACL, #4473]; [BOW: pl. 13, f. 18]
Boloria astarte ; [BNA: 322, #247, pl. 26]; [BOC, 285]; [Opler]

Larva on Saxifraga bronchialis [NAB]

C. a. astarte

NE.British Columbia, Alberta Boloria astarte astarte
Boloria astarte astarte ; [BNA, #247]

Superspecies tritonia. [BARU]

Clossiana tritonia (Böber, 1812)

-. See [About maps] tritonia (Böber, 1812); Mém. Soc. Imp. Nat. Moscou 3: 20, TL: [Lake Baikal, Irkutsk, Russia]
Clossiana tritonia, Lepidoptera collection [SZMN]
Boloria tritonia, NSG Voucher Specimen [Wahlberg]
Clossiana tritonia ; [BARU, #206a]; [BRU2: 51, pl. 34, f. 7-9, 19-21]
Boloria tritonia ; [Otakar Kudrna]

Larva on Saxifraga bronchialis [BARU]

C. t. tritonia

Baikal Clossiana tritonia tritonia ; [BRU2, 51]

C. t. amphilochus (Ménétriés, 1859)

Amur, Ussuri Clossiana tritonia amphilochus ; [BRU2, 51]

Clossiana elatus (Staudinger, 1892)

Transbaikalia. See [About maps] elatus (Staudinger, 1892); , TL: Mongolia, Kentey
Clossiana elatus ; [BARU, #205]
Clossiana tritonia elatus ; [BRU2, 51]

Larva on Saxifraga bronchialis, Vaccinium uligonosum [BARU]

Clossiana distincta (Gibson, 1920)

-. See [About maps] distincta (Gibson, 1920); Rep. Canadian arctic expedition 1913-18. 3 (1): 25(i), 54(i), TL: Harrington Creek, Yukon Territory, Canada: lat 65°05'
Clossiana astarte distincta ; [NACL, #4473a]
Clossiana distincta ; [BARU, #206b]; [BRU2: 51, pl. 34, f. 10-18]

Larva on Saxifraga spinula, (Russia) Salix, (N.America) [BRU2, 51]

C. d. distincta

Yukon - British Columbia

Misidentified/misapplied:

Boloria astarte distincta; [BNA, #247]

C. d. machati Korshunov, 1987

Polar Urals Clossiana distincta machati ; [BARU]; [BRU2, 51]

C. d. dubatolovi Korshunov, 1987

Transbaikalia Clossiana distincta dubatolovi ; [BARU]; [BRU2, 51]

C. d. tschukotkensis (Wyatt, 1961)

NE.Yakutia, Chukotka Clossiana distincta suntara Dubatolov, 1997
Clossiana distincta tschukotkensis ; [BARU]; [BRU2, 51]

Misidentified/misapplied:

?Boloria astarte tschukotkensis; [BNA, #247]

C. d. ershovi Korshunov & Gorbunov, 1995

C.Yakutia Clossiana distincta ershovi Korshunov & Gorbunov, 1995; , TL: Yakutsk
Clossiana distincta ershovi ; [BRU2, 51]

Clossiana matveevi Gorbunov & Korshunov, 1995

Altai. See [About maps] Clossiana matveevi Gorbunov & Korshunov, 1995; [BARU, #206c], TL: Altai, Kurayskiy Mountain rnage, Aktash, 2700m
Clossiana matveevi, Lepidoptera collection [SZMN]
Clossiana matveevi ; [BRU2: 51, pl. 35, f. 10-11]

Clossiana erda (Christoph, 1893)

-. See [About maps] erda (Christoph, 1893); D. Ent. Zeit. Iris, 6: 87, TL: "Vilui" [Vilui R., Yakutia]
Clossiana erda ; [BARU, #203]; [BRU2: 50, pl. 35, f. 4-9]

Larva on Vaccinium vitis-idaea [BRU2, 50]

C. e. erda

Yakutia Clossiana erda erda ; [BRU2, 50]

C. e. kitoica Belik, 1996

E.Sayan Clossiana erda kitoica ; [BRU2, 50]

C. e. puella Churkin, 1999

Transbaikalia (Barguzinsky Mts.) Clossiana erda puella Churkin, 1999; , TL: Transbaikalia (Barguzinsky Mts.)
Clossiana erda puella ; [BRU2, 50]

C. e. dulkeiti Kurenzov, 1970

Magadan Clossiana dulkeiti Kurenzov, 1970; , TL: Omsukchansky Mts., Magadan
Clossiana erda dulkeiti ; [BARU]; [BRU2, 50 (?)]

Clossiana iphigenia (Graeser, 1888)

Japan. See [About maps] iphigenia (Graeser, 1888); Berl. Ent. Ztg. 32: 90, TL: Amur "downstream of Khabarovsk"
Clossiana iphigenia ; [BARU, #191]; [BRU2: 45, pl. 32, f. 10-12]

Larva on Viola selkirkii, (mainland,Japan) Viola grypoceras, (Sakhalin,Japan) [BARU] [BRU2, 45]

C. i. iphigenia

E.Amur, Ussuri, NE.China Clossiana iphigenia iphigenia ; [BRU2, 45]

C. i. alpharatoria Korb, 1997

Sakhalin, Kunashir sachalinensis (Matsumura, 1911); (preocc.)
Clossiana iphigenia sachalinensis ; [BARU]
Clossiana iphigenia alpharatoria ; [BRU2, 45]

19.7.2008 (30)

References:

[About maps]

Warning! The maps are automaticly generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.

[BAMONA]

Paul Opler, Harry Pavulaan, Ray Stanford, Michael Pogue; Butterflies and Moths of North America; Mountain Prairie Information Node

[BARU]

Korshunov, Y. P., Gorbunov, P. Y.; Butterflies of the Asian part of Russia; English translation by Oleg Kosterin; [Dnevnye babochki aziatskoi chasti Rossii. Spravochnik]

[BMAT]

Tennent, John, 1996; The butterflies of Morocco, Algeria and Tunisia; ISBN 0 906802 05 9

[BNA]

Scott, James A., 1986; The Butterflies of North America

[BOA]

Interactive Listing of American Butterflies; Listado Interactivo de la Mariposas Americanas; Butterflies of America

[BOC]

Robert Michael Pyle, 2002; The Butterflies of Cascadia

[BOW]

Lewis, H. L., 1974; Butterflies of the World; ISBN 0 245 52097 X

[BRU2]

Tuzov (ed.), Bogdanov, Churkin, Devyatkin, Dantchenko, Murzin, Samodurov, Zhdanko, 2000; Guide to the Butterflies of Russia and adjacent territories; Libytheidae, Danaidae, Nymphalidae, Riodinidae, Lycaenidae; Volume 2

[EBW]

Smart, Paul, 1976; The Illustrated Encyclopedia of the Butterfly World; ISBN 0 600 31381 6

[HBN]

Francis Hemming, 1937; HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2

[Harri Arkkio]

Finland

[Joseph Belicek]

e-mail: Joseph Belicek

[Kimmo Silvonen]

Espoo, Finland; Kimmo's Lep Site

[LAMAS-2005]

Gerardo Lamas; Entomofauna 26 (6): 57-100; A Bibliography of the Zoological Publications of Hans Fruhstorfer (1886* - 1922+)

[Lars Andersen]

http://www.pedersen-bike.dk/indextre.html

[MBE]

Moths and Butteflies of Europe; Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/

[MRS]

Chou Io (Ed.); Monographia Rhopalocerum Sinensium, 1-2

[Mario Maier]

Europäische Schmetterlinge; e-mail: Mario Maier

[Michael Borsch]

Geesthacht, Germany; contact through; e-mail: Francisco Welter-Schultes

[NAB]

Pyle, R. M.; National Audubon Society: Field Guide to North American Butterflies; 1981; ISBN 0-394-51914-0

[NACL]

Hodges, Ronald W. (ed.), 1983; Check List of the Lepidoptera of America North of Mexico

[NORLEP]

Norges Sommerfugler

[Norbert Kondla]

Canada, BC; e-mail: Norbert Kondla

[Oleg Kosterin]

Institue of Cytology & Genetics; Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/

[Opler]

Opler & Warren, 2003 (April); Butterflies of North America. 2. Scientific Names List for Butterfly Species of North America, north of Mexico.

[Pertti Pakkanen]

e-mail: Pertti Pakkanen; Butterfly and moth observations from Finland

[SPRK]

Seppänen, E. J, 1970; Suomen suurperhostoukkien ravintokasvit, Animalia Fennica 14

[SVF]

Svenska fjärilar; Naturhistoriska riksmuseet; (Sweden)

[SZMN]

Lepidoptera; Siberian Zoological Museum

[Simon Coombes]

1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk

[Wahlberg]

Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group

Some related literature:

Alphéraky, 1881

Lépidoptères du district de Kouldjá et des montagnes environnantes. (1) Horae Soc. ent. Ross. 16 (3-4): 334-435, pl. 14-15

Barnes & Benjamin, 1925

A new subspecies of Brenthis myrina Cram. (Lepid.: Nymphalidae) Ent. News 36: 44

Barnes & McDunnough, 1913

New N. Am. Lepidoptera with notes on described species Contr. nat. Hist. Lep. N. Am. 2 (3): 91-162, pl. 1-9

Butler, 1877

Description of a new species of Argynnis from Arctic America Ent. mon. Mag. 13: 206

Böber, 1812

Contribution de la description de quelques nouvelles espèces de papillons découverts en Sibérie ... Mém. Soc. Imp. Nat. Moscou 3: 20-21,?72

Christoph, 1893

Lepidoptera Nova Faunae Palaearcticae D. Ent. Zeit. Iris, 6: 86-96

Cramer, [1777]

Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 2 (9-16): 1-152, pl. 97-192 (1777)

dos Passos, 1936

Some early stages of Brenthis montinus Scudder Can. Ent. 68: 239-241

Doubleday, [1848]

The genera of diurnal Lepidoptera Gen. diurn. Lep. (1): xii + 1-250, pls. 1-30

Dyar, 1903

A List of North American Lepidoptera and Key to the Literature of this Order of Insects Bull. U.S. natn. Mus., 52: xix, 723pp

Edwards, 1864

Descriptions of certain species of diurnal Lepidoptera, found within the limits of the United States and British America. (3) Proc. ent. Soc. Philad. 2: 501-507

Edwards, 1871

Descriptions of new species of North American butterflies Trans. amer. ent. Soc. 3: 266-277

Edwards, 1890

Description of a new species of Argynnis from Canada [A. alberta] Can. Ent. 22: 113-114

Edwards, 1891

Description of a new species of Argynnis from Alberta Territory Can. Ent. 23: 195-199

Erschoff, 1870

Description d'un Argynnisnouveau de la Sibérie orientale Bull. Soc. Imp. Nat. Moscou 43 (1): 112-114

Eversmann, 1844

De quibsudam Lepidopteria Rossicis Bull. Soc. Imp. Nat. Moscou 17 (3): 588-604, 3pls

Fabricius, 1775

Systema Entomologiae, sistens Insectorum Classes, Ordines, Genera, Species, Adiectis Synonymis, Locis, Descriptionibus, Observationibus. Syst. Ent.: 832pp

Fruhstorfer, 1907

Neue Argynnis Ent. Zs. 21 (27): 163-164 (5 October)

Fruhstorfer, 1908

Neue Argynnis und Melitaea Int. ent. Zs. 1 (41): 310 (11 January)

Fruhstorfer, 1909

Neue palaearktische Rhopaloceren Int. ent. Zs. 3 (4): 20-21 (24 April)

Fruhstorfer, 1909

Neue Palaearkten Int. ent. Zs. 3 (20): 112-113 (14 August)

Gall & Sperling, 1980

A new high altitude species of Boloria from southwestern Colorado, with a discussion of phenetics and hiearchical decisions J. Lep. Soc. 43: 230-252

Gibson, 1920

The Lepidoptera collected by the Canadian arctic expedition 1913-1918. Part I in Report of the Canadian arctic expedition 1918-18. Vol. III, Insects Rep. Canadian arctic expedition 1913-18. 3 (1): 21i-22i

Graeser, 1888

Beiträge zur Kenntnis der Lepidopteren-Fauna des Amurlandes Berl. Ent. Ztg. 32: 33-153 (1888), 309-414 (1889?)

Gunder, 1926

Several new aberrant Lepidoptera (Rhopalocera) Ent. News. 37: 1-9

Gunder, 1928

Additional transitional forms Can. Ent. 60: 162-168, 2pls

Guppy & Kondla, 2002

The correct spellings and type localities of butterflies named from Mount Rainier, Washington, USA J. Lep. Soc. 56 (3): 180-131

Herrich-Schäffer, [1847]

Systematische Bearbeitung der Schmetterlinge von Europa, zugleich als Text, Revision und Supplement zu J. Hübner's Sammlung europäischer Schmetterlinge, 1843 (-1855), Die Tagfalter Syst. Bearb. Schmett. Europ. 1 Text, (ii) Plates _Papilionides_, (iii) Plates _Hesperides_, (9): Title ("1843"), (1): [1-3], 4-16, (ii) pl. 1-5, f. 1-29 (1843), (2): 17-40, (ii) pl. 29-34, f. 131-156 (1843), (3): 41-56, (ii) pl. 6-12, f. 30-60, pl. 35-36, f. 157-164 (1844), (4): 57-80, (ii) pl. 13-22, f. 61-107 (1844), (5): 81-90, (ii) pl. 23-28, f. 108-130, pl. 37-41, f. 165-189 (1844), (6): 91-98, (ii) pl. 42-44, f. 190-203 (1844), (7): 99-114, (ii) pl. 45-52, f. 204-248 (1844), (9): 115-122 (1844), (10): 123-146, (ii) pl. 53-57, f. 249-274 (1845), (11): 147-162 (1845), (12): (iii) pl. 1-3, f. 1-17 (1845), (13): 163-164, (ii) pl. 58-65, f. 275-315 (1846), (14): (ii) pl. 66-70, f. 316-338, (iii) pl. 4-5, f. 18-30 (1846), (26): (ii) pl. 71-77, f. 339-376 (1847), (32): (ii) pl. 78-82, f. 377-400 (1848), (36): (ii) pl. 83-89, f. 401-426, (iii) pl. 6, f. 31-36 (1848), (37): (ii) pl. 90-91, f. 427-437 (1849), (43): (ii) pl. 92-94, f. 438-450, 452-453 (1850), (45): (ii) pl. 95-102, f. 453-489 (1850), (47): (ii) pl. 103, f. 490-495 (1851), (49): (ii) pl. 104-109, f. 496-528 (1851), (51): (ii) pl. 110-118, f. 529-570 (1851), (54): (ii) pl. 119-123, f. 571-596 (1852), (56): (ii) pl. 124-127, f. 597-618, pl. 130-132, f. 628-639, pl. 134, f. 646-651 (1852), (59): (ii) pl. 128-129, f. 619-627, pl. 133, f. 640-645 (1853), (65): (iii) pl. 7, f. 37-42, (index) [1], 2-20 (1854), (66): (index) 21-24 (1855)

Hübner, [1799-1800]; Hübner, [1803-1804]

Sammlung europäischer Schmetterlinge. I. Papiliones - Falter ("Erste Band") Samml. eur. Schmett. [1] (Ziefer?): 3-74, : pl. 1-88, f. 1-446 (1799-1800), pl. 89-96, f. 447-490 (1800-1803), pl. 97-114, f. 491-588 (1803-1804), pl. 115-119, f. 589-612 (1805), pl. 120-124, f. 613-636 (1805-1806), pl. 125-128, f. 637-652 (1806-1808), pl. 129-144, f. 653-727 (1808-1813), pl. 145-147, f. 728-741 (1814-1816), pl. 148-150, f. 742-756 (1816-1817), pl. 151, f. 757-762 (1818-1819), pl. 152-154, f. 757-774 (1819-1822), pl. 155, f. 775-778 (1822-1823), pl. 156-161, f. 779-806 (1823), pl. 162-175, f. 807-874 (1823-1824), pl. 176, f. 869-874 (1824-1825), pl. 177-181, f. 875-895 (1825-1826), (Geyer), pl. 182-187, f. 896-981 (1827-1828), pl. 188-195, f. 922-968 (1828-1832), pl. 196, f. 969-974 (1832-1833), pl. 197-201, f. 975-997 (1834-1836), pl. 202-203, f. 998-1009 (1836-1838), pl. 204-207, f. 1010-1029 (1837-1838)

Kohler, 1977

Revision of North American Boloria selene (Nymphalidae) with description of a new subspecies J. Lep. Soc. 31 (4): 243-268

Linnaeus, 1758

Systema Naturae per Regna Tria Naturae, Secundum Clases, Ordines, Genera, Species, cum Characteribus, Differentiis, Symonymis, Locis. Tomis I. 10th Edition Syst. Nat. (Edn 10) 1: -

Linnaeus, 1767

Systema Naturae per Regna tria Naturae, secundum Classes, Ordines, .... Editio Duocecima Reformata. Tom. 1. Part II. Syst. Nat. (Edn 12) 1 (2): -,533-1327

Sovinsky, 1905

Tri novyk kavkazskikh raznovidnosti cheshuekrylykh Russk. ent. Obozr. 5 (3-4): 108-110

Stallings & Turner, 1947

New American butterflies Can. Entomol. 78 (7-8): 134-135

Staudinger, 1881

Beitrag zur Lepidopterenfauna Central-Asiens Stett. ent. Ztg 42: 253-300, 393-424

Thunberg, 1791

Dissertiom Entomologica sisten Insecta Suecica, ... Diss. Ent. sistens Insecta Suecica (1): 1-24 (1): 1-24 (1 plate) (1784), (2): 32-46 (9 unnumbered) (1791), (3): 48-52 (2 unnumbered) (1792), (4): 53-62 (1792)

Troubridge & Wood, 1990

Biology and taxonomic status of Boloria natazhati (Gibson) J. Lep. Soc. 44 (3): 180-187

Wyatt, 1961

Additions to the Rhopalocera of Afghanistan with descriptions of new species and subspecies J. Lep. Soc. 15 (1): 1-18, 3pls

Zetterstedt, 1839

Insecta Lapponica Insecta Lapponica : 1-1014 ([1839]), 1015-1140 (1840)

If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxononic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.